Michael begins his journey at Siccar Point, near Dunbar on the Firth of Forth, which was the site of an extraordinary discovery in the 18th century. He takes a boat to Bass Rock, before heading to the Scottish capital and Holyrood House.
Michael continues his 300 mile journey along the Firth of Forth at the iconic Forth Bridge, before heading to the port of Leith to tour the Royal Yacht Britannia and then encounters a pair of vast steel sculptures in the Forth Valley.
On this leg of his Scottish railway journey, Michael travels from the medieval city of Stirling through Perth, the historic crowning place of Scottish monarchs, and into Kirkcaldy, where he revisits some fond family memories.
Michael visits St Andrews, home to Scotland's oldest university, in eastern Fife, before heading to the vibrant city of Dundee to learn about the new industry transforming the city and then on to the harbour town of Stonehaven.
Michael Portillo's Scottish railway journey reaches Aberdeen, from which he heads to the wild and beautiful Forvie National Nature Reserve and Scotland's most easterly town, Peterhead.
Michael admires the epic scenery of Cairngorms National Park. He visits the banks of Loch Morlich and learns about writer, poet and naturalist Nan Shepherd, who captured the essence of the Cairngorms in her prose.
Invergordon on the Cromarty Firth, the third great inlet on the northeast coast of the Highlands is visited by Michael Portillo. He explores an ex oil drilling rig now moored in the firth that rather than drilling wells, now caps old ones.
Michael stops at Dunrobin Castle station, which serves the castle, seat of the earls and dukes of Sutherland since the 13th century. He finds out about the second duke of Sutherland and of the Highland Clearances.
Michael crosses over to the Orkney Islands. The former naval base at Scapa Flow is visited and he discovers the islands' extensive Norwegian connections.
One of the destinations Michael visits is Long Sands beach. He helps volunteers caring for sick and injured seals at the Tynemouth Aquarium Seal Hospital such as assisting in taking the temperature of a young pup called Gandalf.
As his journey heads north from Newcastle, Michael discovers Ouseburn which is one of the city's trendiest hotspots. He is intrigued to hear of its part in Newcastle's early industrial development.
A historic collection of Northumbrian bagpipes is awaiting Michael in Morpeth. More practice is needed when he attempts to play one of the wind instruments.
Michael reaches Britain's most northern town, Berwick-upon-Tweed. He joins in a ceremony at the border between Scotland and England on horseback. He rides along the line agreed by the countries in a truce that occurred in 1438.
On this part of his journey Michael investigates the Highland Boundary Fault and how it separates the Highlands from the Lowlands. He also learns how Scotland's geology has influenced its history.
From the fishing port of Oban, Michael travels to the Hebrides. Here, he partakes in seaweed harvesting, pays a visit to the cradle of Christianity in Scotland and sees a cave that was an inspiration for Felix Mendelssohn's music.
Michael crosses the sea to Skye. The island is renowned for wildlife and he seeks out one of its most spectacular residents, the sea eagle while also being lucky enough to spot a school of dolphins.
A new railway journey along the south coast of Wales is begun by Michael Portillo. Along the way, he visits a largely intact Roman amphitheatre and learns about the regeneration efforts in the mining town of Six Bells.
Michael meets a young girl and her family on the Jurassic shoreline. She has a big claim to fame and he learns how her internationally significant discovery on the beach at Barry shook the palaeontology community.
The site of the world's first passenger railway, the Mumbles is visited. Now home to new research into sea walls. New sea defences designs which take both biodiversity and engineering properties into account are being tested.
Michael visits Pembroke and its magnificent Norman castle. He discovers the origins of the Welsh flag and the beginning of one of Britain's greatest royal dynasties - the Tudors. In 1457, Henry VII was born at this castle.
Michael's last destination is Fishguard. He learns the town came under attack by French forces during the 18th century in the last recorded invasion on British soil. A Bayeux-style tapestry commemorating the event is admired by Michael.